Camilla Alves Souto

Assistant Professor of Marine Science

Phone: 609.626.3107
Room: USC2-303


Image of Stockton S logo/Osprey



Ph.D. Integrative Biology, University of California, Berkeley
M.S. Zoology, Universidade Federal da Bahia, Brazil
B.S. Biology, Universidade Federal da Bahia, Brazil


Invertebrate Zoology and Paleobiology, Systematics, Marine Biogeography, Macroevolution


I am interested in understanding the processes that have shaped the evolution of marine life over millions of years. This integrative approach, which intersects Marine Biology and Paleontology, allows me to study broad-scale animal evolution in deep time. I am particularly interested in diversity change, the evolution of body shape, and the processes that dictate the distribution of marine invertebrates in modern and ancient oceans. My research is highly collaborative and focused primarily on echinoderms (sea urchins, sea cucumbers and allies).

I look forward to having students deeply involved in all aspects of my lab’s research, including collection, analysis, and interpretation of data, writing of manuscripts, conference presentation and social media engagement. In addition to my own research interests, I would be thrilled to support dedicated students interested in related research topics.


Martins L, Souto C & Tavares M. 2021. Exploring the macrostructural anatomy of dendrochirotid sea cucumber’s (Echinodermata) calcareous rings under micro-computed tomography (μCT) and its bearing on phylogeny. J Anat, 238(6), 1386-1403.

Cunha R, Martins L, Menegola C, Souto C. 2021. Taxonomy of the sea stars (Echinodermata: Asteroidea) from Bahia State, including ontogenetic variations and an illustrated key to the Brazilian species. Zootaxa, 4955(1), 1–78. 

Martins L & Souto C. 2020. Taxonomy of the Brazilian Apodida (Holothuroidea), with the description of two new genera. Mar Biol Res, 16(4), 219–255.

Souto C, Mooi R, Martins L, Menegola C & Marshall CR. 2019. Homoplasy and extinction: the phylogeny of the cassidulid echinoids (Echinodermata). Zool J Linn Soc, 187, 622–660.

Martins L & Souto C. 2019. Taxonomic remarks on Havelockia and Thyone (Holothuroidea: Dendrochirotida), with descriptions of two new species from the Brazilian coast. JMBA, 99, 1127–1134.

Souto C & Martins L. 2018. Synchrotron micro-CT scanning leads to the discovery of a new genus of morphologically conserved echinoid (Echinodermata: Cassiduloida). Zootaxa, 4457(1), 70–92.

Souto C, Martins L & Menegola C. 2018. Giving up on elaborate dermal ossicles: a new genus of ossicleless Apodida (Holothuroidea). JMBA, 98(7), 1685–1688.

Martins L, Souto C, Braga J & Tavares M. 2018. Echinoidea and Holothuroidea (Echinodermata) of the Trindade & Martin Vaz Archipelago, off Brazil, with new records and remarks on taxonomy and species composition. JMBA, 98(3), 521–555.

Martins L & Souto C. 2015. Taxonomic review of four western Atlantic dendrochirotids (Holothuroidea) with the description of a new Brazilian cucumariid species and designation of neotypes. Zootaxa, 3919(2), 362–374.

Martins L, Souto C, Magalhães WF, Alves OFS, Rosa I & Sampaio CLS. 2012. Echinoderm harvesting in Todos-os-Santos Bay, Bahia state, Brazil: the aquarium trade. Sitientibus, 12, 53–59.

Martins L, Souto C & Menegola C. 2012. A new genus and new species of Sclerodactylidae (Holothuroidea) from the Southwestern Atlantic coast. Zootaxa, 3506, 54–62.

Martins L, Souto C & Menegola C. 2012. First record of Holothuria (Theelothuria) princeps and Thyone pawsoni [Echinodermata: Holothuroidea] in the South Atlantic Ocean. Mar Biodivers Rec, 5, e98.

Souto C, Manso CLC & Martins LR. 2011. Rediscovery and redescription of Cassidulus infidus (Echinoidea: Cassidulidae) from Northeastern Brazil. Zootaxa, 3095, 39–48.